Vegetation Response Following Thinning in Young Douglas-fir Forests of Western Oregon


Book Description

Across western Oregon, Washington, and British Columbia, forest management practices over the past century reduced the amount of late-successional forest while simultaneously increasing the amount of young (less than 80 years old), managed Douglas-fir (Pseudotsuga menziesii) dominated forests. Recently, concerns over loss of late-successional habitat pushed management objectives on public lands away from timber production and toward maintenance and restoration of late-successional habitat. In accordance with these new objectives, The Young Stand Thinning and Diversity Study (YSTDS) was developed to test if thinning could accelerate development of latesuccessional habitat in young managed Douglas-fir forests. Though the YSTDS examines several components of forest ecosystems, the goal of this study was to investigate short-term (5-7 years post-treatment) responses of vegetation to thinning treatments and to evaluate this response in relation to long-term objectives of late-successional development. The study is located on the western slope of the central Oregon Cascades. It consists of four replications of four thinning treatments (treatment areas average 30 ha each) in 30-50 year old second-growth Douglas-fir forest stands. Treatments include a control, heavy thin, light thin, and light thin with gaps. Unlike traditional thinning, the thinning treatments in this study sought to maintain and enhance overstory structural diversity by: (1) retaining species other than Douglas-fir, (2) simulating low densities that characterized development of some old-growth stands, and (3) adding canopy gaps to enhance spatial diversity. Following treatment completion, first, third, and fifth-year vegetation responses were measured Results for overstory vegetation indicate that heavy thinning may accelerate development of large trees, one important component of old-growth structure. This was evident by faster growth of the largest trees in the heavy thin than in the control. A heavy thin may also permit more time for understory development than a lighter thin because canopies of heavy thinned stands remained open longer than canopies of light thinned stands. Variation in overstory cover, which may promote heterogeneous understory development, was higher in the treatment that included canopy gaps than in other treatments including the control. Although accelerated development of a multi-layered canopy was not evident in any treatment, retention of non-dominant tree species prevented simplification of vertical canopy structure by retaining layers that are typically removed by a low thinning prescription. In addition, mortality of non-dominant species was not greater in thinned treatments than in the control. In the understory, results suggest that thinning can increase abundance of some vegetative layers without encouraging homogenization of the understory by clonal shrubs or exotic species. The thinnings resulted in initial declines of bryophytes, tall shrubs, and low shrubs followed by subsequent recovery and growth. While herbs displayed little initial response, a release of early-seral species was evident by 5-7 years post-treatment. Initial changes following thinning were likely due to harvesting damage and/or alteration of microclimate while subsequent changes were probably also related to increased resource availability. It is expected that eventually similarities and differences in overstory structure among thinned treatments will be reflected in the understory. For example, variation in canopy cover created by the addition of canopy gaps was already reflected in the understory, as plant assemblages differed across the gradient from gaps to the thinned forest matrix. Hence, although understory vegetation was similar among heavy and light thins in the short-term, early closure of the canopy following a light thin could preclude continuation of late-seral understory development. Finally, the effect of canopy gaps on the understory was more apparent at a within-stand scale than at a stand scale. Had the within-stand scale been ignored, relevant information regarding understory response would have been overlooked. This indicates that spatial scale should be considered when assessing ecological patterns. In conclusion, it is acknowledged that there are drawbacks to thinning (e.g., certain species decline following thinning) It is also acknowledged that the short-term nature of the data permits only speculation regarding long-term succession. While these limitations are recognized, current trends indicate that a moderate to heavy thinning in combination with gap formation can hasten development of late-successional features in thinned stands relative to unthinned stands. Thus, thinning similar to that used in this study can be one useful tool in the management of young Douglas-fir forests.










Patterns in Understory Vegetation Communities Across Canopy Gaps in Young, Douglas-fir Forests of Western Oregon


Book Description

Canopy gap formation is a major factor contributing to maintenance of overstory species diversity and stand structure in forests and may be integral to development of understory shrub and herb layers as well. Acknowledgement of gap formation as a fundamental feature of natural forests has led to consideration of gaps as an option in forest management regimes. This study examined understory vegetation communities across canopy gaps created as a part of the Density Management Study (DMS), which investigates the effectiveness of a thinning regime in promoting late-successional habitat development in young Douglas-fir forests of western Oregon. Patterns in understory vegetation community composition in and around 0.1 and 0.4ha gaps created as a part of the DMS treatment were investigated. The primary goal of this research was to investigate the potential role of canopy gap creation in fostering heterogeneity in understory vegetation communities, and to examine the extent of gap influence on the surrounding thinned forest matrix. Tree species distributions have been shown to partition across gaps in tropical forest systems through differential responses of species to gradients in resource availability, a pattern known as gap partitioning. In temperate forests, understory vegetation communities are much more diverse than the overstories, and display a greater array of habitat requirements. Therefore, understory communities may be more likely than overstories to exhibit gap partitioning in these forests. Patterns in understory community composition across gaps suggest that gap partitioning has occurred. The strength of this partitioning effect appears to differ between gap sizes, as smaller gaps showed a less powerful effect. Abundance of ruderal species was strongly related to gap partitioning in larger gaps, while smaller gaps were dominated by competitor species. Partitioning may be related to an interactive relationship between harvest-related ground disturbance and resource gradients. Therefore, considerations of gap partitioning processes should take into account intensity and spatial distribution of ground disturbance in relation to resource gradients. In addition, conditions necessary for the expression of gap partitioning in understory vegetation communities may be rare in natural gaps in this region. The influence of gaps on understory vegetation communities in the surrounding forest appears to be relatively small. This small influence extent may help explain the lack of a stand level response to gap formation in these stands. Larger gaps exhibit a slight influence on the understory plant community in the surrounding forest to the north of the gap. In small gaps, there seemed to be an influence of the surrounding forest on gap interiors, resulting in an area of influence smaller than the physical gap area. This relationship may indicate that the area of gap influence on understory vegetation may not scale linearly with physical gap size. Species diversity was higher in gap interiors than in surrounding thinned forests. However this effect was partially due to the presence of exotic species, which showed an affinity for gap interiors. Late successional associated species were negatively related to gap interiors, but only in the larger gap size. Gap creation appears to be promoting small scale species diversity in these stands, but creation of large gaps may also promote the establishment of exotic species and may have a negative effect on late successional associated species. However, any and all of these effects may be transient, as understory communities will be strongly affected by overstory re-establishment, and related changes in resource availability. In general, gap formation may influence small-scale stand heterogeneity as evidenced by understory plant communities, but this effect may rely strongly on the nature of gap formation and intensity of disturbance related to this formation.







Overstory Density and Disturbance Impacts on the Resilience of Coniferous Forests of Western Oregon


Book Description

A trait based approach was used to assess impacts of overstory density and thinning on understory vegetation components related to wildlife habitat. The relationship between overstory basal area and understory vegetation for species grouped by traits, such as production of flowers, fleshy-fruit and palatable leaves, was characterized in thinned and unthinned stands at seven Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) forests in western Oregon six years following harvests. The ranges of overstory densities within thinned and unthinned stands represent gradients of resource availability and thinning disturbance. Lower overstory densities and thinnings were associated with improved ecosystem functions, specifically the provision of wildlife habitat, as evident by higher cover of flowering and fleshy-fruit and palatable leaf producing species. Greater cover of drought, fire and heat tolerant species in low density stands and after thinnings suggested that these ecosystem functions are more likely to be maintained under climate change conditions, indicating higher resilience. The response of specific functions and response types reflect the traits characteristic for each species group and the impact of these traits on sensitivity to resource availability and disturbances. Thus, the correlation between grouping criteria and the main gradients created by management activities can provide an indication of the expected vegetation response, and therefore the impact of management practices on resilience.




Plant Biodiversity


Book Description

Results of regular monitoring of the species diversity and structure of plant communities is used by conservation biologists to help understand impacts of perturbations caused by humans and other environmental factors on ecosystems worldwide. Changes in plant communities can, for example, be a reflection of increased levels of pollution, a response to long-term climate change, or the result of shifts in land-use practices by the human population. This book presents a series of essays on the application of plant biodiversity monitoring and assessment to help prevent species extinction, ecosystem collapse, and solve problems in biodiversity conservation. It has been written by a large international team of researchers and uses case studies and examples from all over the world, and from a broad range of terrestrial and aquatic ecosystems. The book is aimed at any graduate students and researchers with a strong interest in plant biodiversity monitoring and assessment, plant community ecology, biodiversity conservation, and the environmental impacts of human activities on ecosystems.










Integrated Restoration of Forested Ecosystems to Achieve Multiresource Benefits


Book Description

"A primary mission of the U.S. Department of Agriculture Forest Service is multiple resource management, and one of the emerging themes is forest restoration. The National Silviculture Workshop, a biennial event co-sponsored by the Forest Service, was held May 7-10, 2007, in Ketchikan, Alaska, with the theme of "Integrated Restoration of Forested Ecosystems to Achieve Multiresource Benefits." This proceedings presents a compilation of state-of-the-art silvicultural research and forestry management papers that demonstrates integrated restoration to yield multiple resource benefits. These papers highlight national perspectives on ecosystem services, forest restoration and climate change, and regional perspectives on forest restoration and silvicultural practices to achieve multiple resource benefits from researchers and forest practitioners working in a broad array of forest types in the United States."